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Movements comprise both dynamic (displacements) and static (successive positions) parameters that are encoded into phasic and tonic discharge, the latter playing also a role as an internal reference. Combining the two components allows a rapid (phasic) and referenced (tonic) spatial representation of joint position and movement. In the crayfish, phasic coding is a property of all CBCO sensory neurons, and their firing frequency during actual movement is always higher than during stationary plateaus, indicating that the major information conveyed by sensory afferents is dynamic rather than static.
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